| Date | Filer | Title | Filing | Type | Shares Traded | Price | Total Held |
|---|---|---|---|---|---|---|---|
| Aug 23, 2018 |
Ten Percent Owner
Trans History: 20
|
Ten Percent Owner | Form 4 | Conversion of derivative security | 3,682,919 | -- | -- |
| Aug 23, 2018 |
Chief Scientific Officer
Trans History: 21
|
Chief Scientific Officer | Form 4 | Conversion of derivative security | 4,362,062 | -- | -- |
| Aug 23, 2018 |
Ten Percent Owner
Trans History: 20
|
Ten Percent Owner | Form 4 | Conversion of derivative security | 5,426,808 | -- | -- |
| Aug 23, 2018 |
Ten Percent Owner
Trans History: 20
|
Ten Percent Owner | Form 4 | Conversion of derivative security | 6,396,105 | -- | -- |
| Aug 23, 2018 |
Ten Percent Owner
Trans History: 12
|
Ten Percent Owner | Form 4 | Conversion of derivative security | 6,443,565 | -- | -- |
| Aug 23, 2018 |
Director, Ten Percent Owner
Trans History: 14
|
Director, Ten Percent Owner | Form 4 | Conversion of derivative security | 6,443,565 | -- | -- |
| Aug 23, 2018 |
Director, Ten Percent Owner
Trans History: 14
|
Director, Ten Percent Owner | Form 4 | Conversion of derivative security | 6,443,565 | -- | -- |
| Aug 23, 2018 |
Ten Percent Owner
Trans History: 20
|
Ten Percent Owner | Form 4 | Conversion of derivative security | 6,810,098 | -- | -- |
| Aug 23, 2018 |
Ten Percent Owner
Trans History: 12
|
Ten Percent Owner | Form 4 | Conversion of derivative security | 8,545,184 | -- | -- |
| Aug 23, 2018 |
Director, Ten Percent Owner
Trans History: 14
|
Director, Ten Percent Owner | Form 4 | Conversion of derivative security | 8,545,184 | -- | -- |
* Represents the number of non-derivative securities directly held. (I) Indicates that some (or all) of these shares are held indirectly.