Date Filer Filing Type Shares Traded Price Total Held
Oct 01, 2018
Chief Scientific Officer
Form 4 Grant, award or other acquisition pursuant to Rule 16b-3(d) 23,230 -- 23,230
Aug 23, 2018
Chief Scientific Officer
Form 4 Conversion of derivative security 5,941 -- 7,354
Aug 23, 2018
Chief Scientific Officer
Form 4 Conversion of derivative security 1,231 -- 8,585
Aug 23, 2018
Chief Scientific Officer
Form 4 Conversion of derivative security 101,918 -- 110,503
Aug 23, 2018
Chief Scientific Officer
Form 4 Conversion of derivative security 2,433 -- 112,936
Aug 23, 2018
Chief Scientific Officer
Form 4 Conversion of derivative security 254,275 -- --
Aug 23, 2018
Chief Scientific Officer
Form 4 Conversion of derivative security 52,720 -- --
Aug 23, 2018
Chief Scientific Officer
Form 4 Conversion of derivative security 4,362,062 -- --
Aug 23, 2018
Chief Scientific Officer
Form 4 Conversion of derivative security 104,167 -- --
Aug 21, 2018
Chief Scientific Officer
Form 3 Initial Statement of Beneficial Ownership of Securities -- -- 1,413

* Represents the number of non-derivative securities directly held. (I) Indicates that some (or all) of these shares are held indirectly.