Date Filer Title Filing Type Shares Traded Price Total Held
Jan 18, 2019
Chief Operating Officer
Chief Operating Officer Form 4 Exercise or conversion of derivative security exempted pursuant to Rule 16b-3 6,000 $1.30 6,815
Jan 18, 2019
Chief Operating Officer
Chief Operating Officer Form 4 Exercise or conversion of derivative security exempted pursuant to Rule 16b-3 6,000 -- 28,219
Aug 23, 2018
Chief Scientific Officer
Chief Scientific Officer Form 4 Conversion of derivative security 5,941 -- 7,354
Aug 23, 2018
Chief Scientific Officer
Chief Scientific Officer Form 4 Conversion of derivative security 1,231 -- 8,585
Aug 23, 2018
Chief Scientific Officer
Chief Scientific Officer Form 4 Conversion of derivative security 101,918 -- 110,503
Aug 23, 2018
Chief Scientific Officer
Chief Scientific Officer Form 4 Conversion of derivative security 2,433 -- 112,936
Aug 23, 2018
Chief Scientific Officer
Chief Scientific Officer Form 4 Conversion of derivative security 254,275 -- --
Aug 23, 2018
Chief Scientific Officer
Chief Scientific Officer Form 4 Conversion of derivative security 52,720 -- --
Aug 23, 2018
Chief Scientific Officer
Chief Scientific Officer Form 4 Conversion of derivative security 4,362,062 -- --
Aug 23, 2018
Chief Scientific Officer
Chief Scientific Officer Form 4 Conversion of derivative security 104,167 -- --

* Represents the number of non-derivative securities directly held. (I) Indicates that some (or all) of these shares are held indirectly.