Date Filer Title Filing Type Shares Traded Price Total Held
Oct 01, 2018
Director, Ten Percent Owner
Director, Ten Percent Owner Form 4 Grant, award or other acquisition pursuant to Rule 16b-3(d) 10,516 -- 10,516
Oct 01, 2018
Chief Financial Officer
Chief Financial Officer Form 4 Grant, award or other acquisition pursuant to Rule 16b-3(d) 59,590 -- 59,590
Oct 01, 2018
Director, Ten Percent Owner
Director, Ten Percent Owner Form 4 Grant, award or other acquisition pursuant to Rule 16b-3(d) 10,516 -- 10,516
Aug 23, 2018
Chief Operating Officer
Chief Operating Officer Form 4 Open market or private purchase of non-derivative or derivative security 815 $6.12 815
Aug 23, 2018
Chief Operating Officer
Chief Operating Officer Form 4 Open market or private purchase of non-derivative or derivative security 815 -- 815
Aug 23, 2018
Chief Scientific Officer
Chief Scientific Officer Form 4 Conversion of derivative security 5,941 -- 7,354
Aug 23, 2018
Chief Scientific Officer
Chief Scientific Officer Form 4 Conversion of derivative security 1,231 -- 8,585
Aug 23, 2018
Chief Scientific Officer
Chief Scientific Officer Form 4 Conversion of derivative security 101,918 -- 110,503
Aug 23, 2018
Chief Scientific Officer
Chief Scientific Officer Form 4 Conversion of derivative security 2,433 -- 112,936
Aug 23, 2018
Chief Scientific Officer
Chief Scientific Officer Form 4 Conversion of derivative security 254,275 -- --

* Represents the number of non-derivative securities directly held. (I) Indicates that some (or all) of these shares are held indirectly.