Date Filer Title Filing Type Shares Traded Price Total Held
Aug 21, 2018
Director
Director Form 3 Initial Statement of Beneficial Ownership of Securities -- -- --
Aug 21, 2018
Director
Director Form 3 Initial Statement of Beneficial Ownership of Securities -- -- --
Aug 21, 2018
Chief Operating Officer
Chief Operating Officer Form 3 Initial Statement of Beneficial Ownership of Securities -- -- --
Aug 21, 2018
Chief Operating Officer
Chief Operating Officer Form 3 Initial Statement of Beneficial Ownership of Securities -- -- --
Aug 21, 2018
Chief Operating Officer
Chief Operating Officer Form 3 Initial Statement of Beneficial Ownership of Securities -- -- --
Aug 21, 2018
Chief Scientific Officer
Chief Scientific Officer Form 3 Initial Statement of Beneficial Ownership of Securities -- -- 1,413
Aug 21, 2018
Chief Scientific Officer
Chief Scientific Officer Form 3 Initial Statement of Beneficial Ownership of Securities -- -- --
Aug 21, 2018
Chief Scientific Officer
Chief Scientific Officer Form 3 Initial Statement of Beneficial Ownership of Securities -- -- --
Aug 21, 2018
Chief Scientific Officer
Chief Scientific Officer Form 3 Initial Statement of Beneficial Ownership of Securities -- -- --
Aug 21, 2018
Chief Scientific Officer
Chief Scientific Officer Form 3 Initial Statement of Beneficial Ownership of Securities -- -- --

* Represents the number of non-derivative securities directly held. (I) Indicates that some (or all) of these shares are held indirectly.