Date Filer Title Filing Type Shares Traded Price Total Held
Aug 21, 2018
Chief Operating Officer
Chief Operating Officer Form 3 Initial Statement of Beneficial Ownership of Securities -- -- --
Aug 21, 2018
Chief Operating Officer
Chief Operating Officer Form 3 Initial Statement of Beneficial Ownership of Securities -- -- --
Aug 21, 2018
Chief Operating Officer
Chief Operating Officer Form 3 Initial Statement of Beneficial Ownership of Securities -- -- --
Jun 06, 2019
Director
Director Form 4 Grant, award or other acquisition pursuant to Rule 16b-3(d) 21,911 -- 21,911
Oct 01, 2018
Director
Director Form 4 Grant, award or other acquisition pursuant to Rule 16b-3(d) 10,516 -- 10,516
Oct 01, 2018
Chief Scientific Officer
Chief Scientific Officer Form 4 Grant, award or other acquisition pursuant to Rule 16b-3(d) 23,230 -- 23,230
Aug 23, 2018
Chief Scientific Officer
Chief Scientific Officer Form 4 Conversion of derivative security 5,941 -- 7,354
Aug 23, 2018
Chief Scientific Officer
Chief Scientific Officer Form 4 Conversion of derivative security 1,231 -- 8,585
Aug 23, 2018
Chief Scientific Officer
Chief Scientific Officer Form 4 Conversion of derivative security 101,918 -- 110,503
Aug 23, 2018
Chief Scientific Officer
Chief Scientific Officer Form 4 Conversion of derivative security 2,433 -- 112,936

* Represents the number of non-derivative securities directly held. (I) Indicates that some (or all) of these shares are held indirectly.